Vergne is bidding to follow in the wheeltracks of fellow Red Bull Junior Team members Jaime Alguersuari and Daniel Ricciardo and race off with Carlin's third British F3 title in as many seasons.
Leading the charge from the rival teams, 17 points behind Vergne, is Fortec's impressive Oliver Webb, who was a solid runner-up to Vergne in rounds one and three, but was outpaced at the test by team-mate Daniel McKenzie.
There will be a number of drivers at Silverstone hoping to put shaky seasonal starts behind them, not least Racing Steps Foundation-backed man Calado, who suffered car trouble in qualifying at Oulton which blighted his weekend. Another Brit, the 2008 British F3 National champion Jay Bridger, showed top-five pace in qualifying in his Litespeed-prepared Dallara, but went off in race one while holding a safe seventh.
Hitech Racing took comfort from promising showings for Buller and for his team-mate Gabriel Dias at Oulton, the Brazilian bouncing back from adversity to take fourth place in round three.
T-Sport have a one-two stranglehold on the National Class thanks to class victories at Oulton for Idafar and Cole, while the squad's sole International runner, Alex Brundle, is enjoying a solid start to his F3 career.
There's no shortage of British talent on the grid this season, with CF Racing with Manor's talented Welsh driver Hywel Lloyd looking to overcome his embarrassing faux pas
at Oulton, and the contingent will be swelled by the arrival of Euroseries regular Alexander Sims, who, along with two of his colleagues from France's ART Grand Prix team - Mexican Esteban Gutierrez and French driver Jim Pla - will be taking chance to measure his pace against the team's British F3 rivals.